Fifty Years of Darwinism : Modern Aspects of Evolution ; Centennial Addresses in Honor of Charles Darwin, Before the American Association for the Advancement of Science, Baltimore, Friday, January 1, 1909

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Fresh-water fishes are not a group different from salt-water fishes. Many salt-water fishes can enter fresh water, and we may for present purposes assume that all candidates for fresh- "water existence are adapted or readily adaptable to the fresh water. Adaptations to the second and third of the fresh-water conditions imply pe- culiarities in habit or structure not possessed by all fishes, and these must, in the main, have been acquired by the marine fishes before they could enter and maintain
... themselves in fresh water.
ADAPTATION 185 The downward current and sediment, if the lat- ter is not too abundant, are not obstacles suf- ficient to keep an adult fish from entering fresh waters. The eggs and young furnish the point of attack. Among oceanic fishes we have many that have pelagic eggs, others that have adhesive eggs, others that have heavy cohesive eggs, others that have filaments for the attachment of eggs, while others harbor their eggs.
Currents would naturally tend to carry pelagic eggs into the ocean, and as far as I know only one fish with pelagic eggs has succeeded in estab- lishing itself in fresh water, and it, the eel, to the present day, descends to the sea to deposit its eggs!


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