The Relative Value of the Processes Causing Evolution

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Whenever we want to discuss this law and its universal va- lidity, we must remember that in its simplest form, as we have formulated it, it only states that the genes themselves are qualitatively stable. The usual ineffectiveness of selection in pure lines and pure clones for instance, we explain by assuming that in such material the individuals really have all the same set of genes. But at the same time we know, that the ineffect- iveness of selection and of change of environment to change suc
...h material, is only the result of this nature of the genes, and will hold good only so long as within this material there do not occur processes, which change the set of genes carried. And we must not confound a temporary change of the quantitative increase or decrease of genes in the cytoplasing a number of cell-generations with the qualitative change in the genes them- selves.
We may certainly not turn round the statement that genes are qualitatively stable, and that therefore, the germ-plasm of material pure for its set of genes is not amenable to change by selection, and make it read so, that the test of the effectiveness of selection in a pure line or a pure clone is a test of the validity or otherwise, of Johannsen's law.


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