The Elements of Embryology

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The two masses behind the auditory vesicle represent the glossopharyngeal and pneumogastric nerves (Fig. 45, VIII. , Fig. 46, G. Ph. And Pg. }. At first united, they subsequently become separate. The glossopharyngeal supplies the third arch, and the pneumogastric the fourth arch.
These four masses, representing four important mixed cranial nerves, seem to be derived directly from the mesoblast surrounding the hind-brain. It is worthy of notice that they are mixed, sensory and motor, nerves; for
..., restricted as are the sensory functions of the seventh, and the motor func- tions of the pneum. Ogastric in the adult mammal, the study of their com- parative physiology leaves no doubt as to the essentially mixed nature of each. It is also worthy of note that of the third, fourth and sixth nerves, no such early rudiments appear ; and there are reasons for thinking that these are in reality intercranial branches, the third and fourth of the fifth, and the sixth of the seventh nerves. The purely sensory nerve or rather sense-nerve, the auditory, seems to have a different origin altogether from all the above, though it may perhaps be looked upon as the dorsal branch of the seventh, while the erratic hypoglossal appears to be distinctly a spinal nerve.

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